They were the nymphs of evening and the golden light of sunsets.
They were the daughters of either Nyx or the Titan Atlas. The Hesperides were entrusted with the care of the tree of the golden apples, which was had been presented to the goddess Hera by Gaia on her wedding day. They were assisted by Dracon. Heracles was sent to fetch the apples as one of his Twelve Labours and, upon slaying the serpent, stole the precious fruit. Athena later returned them to the Hesperides.